The BoomTown system uses tracking email addresses in the database to capture emails going to your leads, and replies from your leads so that the system can ensure the emails are routed to the correct lead profile, and alert the correct user that's associated with the lead!
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Each lead has a unique tracking email address created automatically, and each Agent or Lender is also assigned a unique tracking email address in association with every one of their leads. The tracking email addresses follow a similar format for each user and will look similar to the image below.

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The tracking email address is created when a user profile or lead profile is created, and for users specifically, the prefix will be the Alias associated with their account, which is the original Agent subdomain that was entered for the user.
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The tracking email addresses are masked with the login email address on a user's account, so in most cases, you and the lead see only the login email address in correspondence.
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For example, if you have a lead named Joe Smith and Joe's email address is joe@gmail.com, BoomTown assigns a tracking email address like 295908420_joe@gmail.com. When you email Joe through the system, you are actually sending an email to 295908420_joe@gmail.com which gets routed to joe@gmail.com. The email you send Joe through BoomTown will be logged in Joe Smith's History in his lead profile.
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If you were to send an email to Joe@gmail.com directly through another email client, like Gmail or Outlook, you will notice that the email will not be captured by the BoomTown system and logged in your lead's History.

If you hit reply to an email that your lead sent you through BoomTown from within your email client, your email response should be logged in the lead's Summary when you are replying to 295908420_joe@gmail.com rather than joe@gmail.com.
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That said, it's best not to get into the habit of emailing your leads from an email client or webmail, since you may not be able to easily tell if you are emailing the tracking link address or the regular address, especially if your lead is saved as a contact and you only see that you are emailing "Joe Smith".