If a response you've sent is in the outbox with this error message, it's due to an attempt of using a template with auto-insert variables for a location that has yet to be assigned those in the application.
Resolution:
Assign the auto-reply aliases to the location(s),
Delete the previous response from the Outbox,
Use the auto-reply template to respond to review.
If done correctly, then after the set delay, your response should successfully post.
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